A retired dentist with a keen interest in palaeontology and the environment has left money from his £1.7m estate to ecological organ- isations.

William Graham-Smith, of White Barn Drive, Boar's Hill near Oxford, died last year shortly before his 90th birthday.

The Friends of the Earth Trust, Greenpeace Environmental Trust and the New Economics Foundation, have all benefited in his will, the probate for which was granted to Franklins Solicitors of Walton House, Ock Street, Abingdon.

A spokesman for the firm said that Mr Graham-Smith and his late wife had no children. She said: "He had a lifelong interest in palaeontology and even write a book on the subject."

Before his death, Mr Graham-Smith left his entire collection of fossil fish and other fossil materials from the Devonian period, plus notes and photographs, to the National Museum of Scotland in Edinburgh.